Used in biochemical research to study enzyme mechanisms, particularly those involving nucleotide metabolism. It serves as a substrate for synthesizing important biomolecules like phospholipids and glycoproteins. Applied in cell culture studies to investigate nucleotide incorporation and cellular signaling pathways. Also utilized in the production of nucleotide-based pharmaceuticals and as a precursor for synthesizing modified nucleotides in molecular biology applications. Plays a role in metabolic pathway analysis and the development of therapeutic agents targeting nucleotide-dependent processes.