Used primarily in scientific research as a specific inhibitor of the sarcoplasmic/endoplasmic reticulum calcium ATPase (SERCA) pump. It helps investigate calcium signaling pathways in cells, particularly in muscle and nerve tissues. Due to its ability to disrupt intracellular calcium homeostasis, it is also used to study cellular stress responses, apoptosis, and endoplasmic reticulum function. Not used in clinical or industrial applications because of its toxicity.